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Glossary of Astrobiology Terms

ALH84001 - Martian meteorite discovered in the Allan Hills ice field in Antarctica in 1984. It has been suggested that this meteorite contains evidence of martian microbial life.

Amino Acid - Simple organic molecules containing an amino group (NH2) and a carboxyl group (COOH). Amino acids link together in chains to form proteins.

Archaea - One of three domains of life, along with the Bacteria and the Eucarya. The archaea are prokaryotic organisms which were shown to be a distinct group from the bacteria by molecular phylogeny using 16S ribosomal RNA. The archaeal domain includes many extremophilic organisms.

Archaebacteria - An alternative name for the domain archaea.

Archean - The period in the history of the Earth from 4000 million years ago to 2500 million years ago, part of the Precambrian. The Archean is preceeded by the Hadean and followed by the Proterozoic. The earliest fossil evidence for life on Earth appears during the Archean period.

ATP - Adenosine triphosphate - The molecule which is the source of energy for most metabolic processe in living organisms.

Bacteria - One of three domains of life, along with the Archaea and the Eucarya. The bacterial domain includes all prokaryotic organisms not classified as archaea.

Carbonaceous Chondrite - A rare type of meteorite which is rich in organic material. The Murchison meteorite is a famous example.

Cenancestor - An alternative term for the Last Common Ancestor of all life on Earth.

Chirality - A chiral molecule is a molecule with an asymmetric structure which can exist in two mirror image forms (or enantiomers). In living organisms such molecules are usually found in only one of the two possible enantiomers (homochirality). Thus amino acids are normally in the left-handed or L-enantiomer, while sugars are in the right-handed or D enantiomer.

Chloroplast - An organelle found in the cells of green plants in which photosynthesis occurs. According to the endosymbiosis theory, chloroplasts are descended from photosynthetic bacteria.

Cyanobacteria - A class of bacteria which make use of oxygen producing photosynthesis. Commonly referred to as blue-green algae.

DNA - Deoxyribonucleic Acid - The molecule which constitutes the genome of living cells. DNA molecules have a double stranded helical structure built from a sugar phosphate backbone and a set of four bases (adenine, guanine, cytosine and thymine). The sequence of bases specifies the genetic information.

Domain - The highest taxonomic division in the classification of living organisms. The three domains are the Archaea, the Bacteria and the Eucarya. Domains are subdivided into kingdoms. While the three domain model is widely used in astrobiology, some biologists prefer other schemes such as the Five-Kingdom system.

Doppler Shift - The shift in wavelength of light or other electromagnetic radiation due to the motion of the source relative to the observer. If the source is moving away from the observer the wavelength of light is increased (a red shift) whereas for a source moving towards the observer the wavelength is decreased (a blue shift). By precise measurements of the observed wavelength of spectral lines astronomers can determine the velocity of a star with accuracies of a few metres per seconds. Such observations are used to detect extrasolar planets.

Drake Equation - An equation proposed by SETI pioneer Frank Drake to calculate the number of civilizations in the galaxy. It takes the form:

N = R fp ne fl fi fc L

where R is the birthrate of suitably long lived stars, fp is the fraction of stars with planets, ne is the number of earth-like planets per solar system, fl is the fraction of earth-like planets on which life actually starts, fi is the fraction of planets with life on which intelligence emerges, fc is the fraction of intelligent civilizations which have the technology and incentive to communicate over interstellar distances, and L is the lifetime of such technological civilizations.

Ediacaran Fauna - A fossil fauna of soft bodied organisms with an age of about 560 million years, first found in the Flinders Ranges of South Australia. They are the earliest known animals.

Enantiomer - The name given to the two mirror image forms of a chiral molecule.

Endosymbiosis - Symbiosis in which one organism lives within the body of the other. More specifically refers to the theory that eukaryotic cells arose from the symbiosis of a number of organisms, with the organelles such as mitochondria and chloroplasts being descended from bacteria. This theory has been confirmed by molecular phylogeny.

Enzyme - A protein that acts as a catalyst. Most chemical processes in living cells are enzyme catalysed.

Eubacteria - An alternative name for the domain bacteria (or true bacteria) as distinct from the archaebacteria or archaea.

Eucarya (or Eukarya) - One of three domains of life along with the Archaea and the Bacteria. The Eucarya domain covers all organisms with cells which have a nucleus and organelles (Eukaryotic cells). The Eucarya domain includes the animals, plants, fungi and protists.

Eukaryote - An organism belonging to the domain Eucarya. Eukaryotes have cells with the genetic material contained in a nucleus and with separate organelles. Eukaryotic cells are typically about 10 times bigger in each dimension than prokaryotic cells.

Extrasolar Planet - A planet orbiting a star other than the sun.

Extremophile - An organism adapted to life in extreme conditions. For example thermophiles and hyperthermophiles are adpated to high temperatures.

Gas Giant - A massive planet such as Jupiter or Saturn composed mainly of gases with no solid surface.

Genetic Code - The set of rules by which three letter "words" in a DNA or RNA sequence describe an amino acid to be incorporated into a protein.

Genome - The complete set of genetic information for a particular organism.

Gravitational Lensing - The effect on the appearance of an astronomical source due to the bending of light by the gravitational field of an intervening object.

Habitable Zone - The region around a star in which an orbiting planet could maintain conditions suitable for life.

Hadean - The period from the formation of the Earth (about 4550 million years ago) to 4000 million years ago. The first division of the Precambrian. The Hadean period is followed by the Archean.

Heavy Bombardment - During the first few hundred million years of the solar system the earth and other planets were subject to an intense bombardment by the debris left over from the formation of the solar system. It is during this heavy bombardment phase that most of the craters on the moon were formed. The emergence of life on Earth appears to roughly coincide with the end of the heavy bombardment.

Hot Jupiter - A massive planet orbiting very close to a star, such as the planet of 51 Pegasi with an orbital period of only four days. Many of the extrasolar planets so far found are of this type.

Hyperthermophile - An organism adapted to life at very high temperatures. Hyperthermophiles have optimum growth temperatures above 80 degrees C, and a number can grow at temperatures above 100 degrees C.

Impact Frustration - The hypothesis that during the heavy bombardment phase, impacts of massive objects may have wiped out emerging life several times before it was able to permanently establish itself on Earth.

Interferometer - An instrument which makes use of the phenomenon of interference of light. Astronomical interferometers work by combining the light of two or more telescopes to achieve the resolving power equivalent to a much larger telescope. Interferometers are widely used in radio astronomy, but the technique is more difficult to apply in the optical because of the disturbing effects of the earth's atmosphere. Space interferometers avoid these difficulties and may play an important role in the search for extrasolar planets.

Interstellar Molecule - Molecules in interstellar space are most commonly detected by means of radio frequency emission lines coming from the gas in molecular clouds. Molecules can also be detected from the infrared spectra emitted from dust. Well over a hundred molecular species have been detected by these methods. Some of the more complex molecules found include acetic acid, acetone and ethanol.

Last Common Ancestor - The last common ancestor of all organisms living today. The root of the tree of life.

Lateral Gene Transfer - The transfer of genes between different species. Lateral gene transfer may have been widespread in the early stages of life on Earth and this complicates the interpretation of the tree of life.

LUCA - Another term used for the Last Common Ancestor of all living organisms. Acronym for Last Universal Common Ancestor.

Martian Meteorites - Meteorites which originate from Mars, also known as SNC meteorites. Their martian origin is demonstrated by bubbles of gas trapped within them which have identical composition to the atmosphere of Mars.

Meteorite - A meteor that reaches the earth's surface. There are many types of meteorites, the most important for astrobiology being the carbonaceous chondrites which contain organic material, and the martian meteorites.

Microlensing - Gravitational lensing of a star indicated by the brightening of the star as the lensing object moves in front of it. Analysis of the light curve of a microlensing event can reveal the presence of an extrasolar planet.

Miller-Urey Experiment - An experiment carried out by Stanley Miller and Harold Urey in 1952 which demonstrated the synthesis of amino acids in conditions simulating a thunderstorm on the early Earth. The experiment can be seen as marking the beginning of the experimental study of the chemistry of life's origins.

Mitochondria - The organelles in eukaryotic cells in which aerobic respiration takes place, providing a supply of chemical energy (in the form of ATP) for the cell.

Molecular Cloud - A cloud of cool gas in interstellar space. These clouds provide the raw material out of which new stars form and contain many types of interstellar molecules.

Nucleic Acid - The molecules which carry genetic information, DNA and RNA.

Organelle - Membrane enclosed structures in eukaryotic cells such as the mitochondria and chloroplasts.

Panspermia - The hypothesis that life has spread from planet to planet throughout the galaxy and therefore did not originate on Earth. The term originates with Svante Arrhenius in the 1900s. More recently the most prominent supporters of this view have been Fred Hoyle and Chandra Wickramasinghe. Most scientists find no evidence to support panspermia and think it unlikely that life could survive a journey between stars. There is somewhat more support for the idea that life could have spread between planets within the solar system (planetary panspermia)

Peptide - A compound linked together by peptide bonds such as a protein.

Photosynthesis - The chemical process carried out in photosynthetic bacteria and green plants in which light energy is used to synthesize carbohydrates from water and carbon dioxide.

Phylogeny - The evolutionary relationships between different species of organisms as represented in a phylogenetic tree. In molecular phylogeny these relationships are determined by analysis of the differences in the sequences of genes common to the various species.

Planetary Panspermia - The spreading of life between planets within the solar system. In particular the hypothesis that life might have originated on Mars and been carried to Earth - a suggestion which is supported by the existence of martian meteorites.

Precambrian - The period in Earth's history from its formation up to the beginning of the Cambrian period about 540 million years ago. The Precambrian is subdivided into the Hadean, the Archean, and the Proterozoic.

Pre-RNA World - A hypothetical early stage in the development of life which preceeded the RNA World and used some other genetic material in place of RNA or DNA.

Prokaryote - An organism belonging to the domains archaea or bacteria. Prokaryotic cells lack a nucleus and organelles and are typically much smaller than eukaryotic cells.

Protein - A polymer built from amino acid monomers. Living organisms use many different proteins acting as catalysts (enzymes) and for structural and other roles.

Proterozoic - The period in Earth's history from 2500 million to 540 million years ago. The last division of the Precambrian.

Ribosome - A structure composed of protein and RNA molecules which reads genetic information from messenger RNA and synthesises the corresponding protein.

Ribosomal RNA - The RNA components of a ribosome. One of these components, the small sub-unit ribosomal RNA (also known as 16S ribosomal RNA in prokaryotes or 18S ribosomal RNA in eukaryotes) has been widely used to determine the tree of life.

Ribozyme - An RNA molecule which acts as a catalyst. The discovery of RNA catalysis led to a Nobel prize for Sidney Altman and Thomas Cech, and to the RNA World concept.

RNA - Ribonucleic acid - A molecule which can carry genetic information in a similar way to DNA. In modern cells RNA molecules are important in the process of protein synthesis, in the form of messenger RNA, ribosomal RNA and transfer RNA.

RNA World - A hypothetical early stage in the development of life in which RNA molecules provided both the genome and the catalysts, roles which subsequently were taken over by DNA and proteins.

SETI - The Search for Extraterrestrial Intelligence.SETI projects aim to detect radio or other signals from advanced civilizations elsewhere in the galaxy.

SNC meteorites - Another term for martian meteorites. The name derives from the initials of three examples, Shergotty, Nakhla and Chassigny.

Snowball Earth - Name given to events in which the Earth was glaciated and the oceans frozen from pole to pole. Snowball Earth events have been suggested as occuring at about 2.3 billion years ago and between 750 and 580 million years ago.

Stromatolite - Layered structures built by colonies of microorganisms which are commonly found in the Archean and Proterozoic fossil records. Modern examples can be found in sites such as Shark Bay in Western Australia.

Terrestrial Planet - Rocky planets such as the Earth and other inner planets, as distinct from the Gas Giants.

Thermophile - An organism adapted to life at high temperatures.

Tree of Life - A phylogenetic tree covering all groups of life on Earth. The term is commonly used for the tree derived by molecular phylogeny using small sub-unit Ribosomal RNA as pioneered by Carl Woese in the 1970s.

   
Jeremy Bailey (jab@aaoepp.aao.gov.au)